The scientific study of human evolution encompasses the development of the genus Homo, but usually involves studying other hominids and hominines as well, such as Australopithecus. "Modern humans" are defined as the Homo sapiens species, of which the only extant subspecies is Homo sapiens sapiens; Homo sapiens idaltu (roughly translated as "elder wise human"), the other known subspecies, is extinct.[5] Anatomically modern humans appear in the fossil record in Africa about 130,000 years ago.[6][7]
The closest living relatives of Homo sapiens are two distinct species of the genus Pan: the Bonobo (Pan paniscus) and the Common Chimpanzee (Pan troglodytes). These species share the same common ancestor. The main difference between them is the social organization: matriarchal for the Bonobo and patriarchal for the Common Chimpanzee. Full genome sequencing resulted in the conclusion that "after 6.5 million years of separate evolution, the differences between bonobo/chimpanzee and human are just 10 times greater than those between two unrelated people and 10 times less than those between rats and mice". In fact, 95 per cent of the DNA sequence is identical between the two Pan species and human.[8][9][10][11] It has been estimated that the human lineage diverged from that of chimpanzees about five million years ago, and from gorillas about eight million years ago. However, a hominid skull discovered in Chad in 2001, classified as Sahelanthropus tchadensis, is approximately seven million years old, which may indicate an earlier divergence.[12]
There are two prominent scientific theories of the origins of contemporary humans. They concern the relationship between modern humans and other hominids. The single-origin, or "out-of-Africa", hypothesis proposes that modern humans evolved in Africa and later migrated outwards to replace hominids in other parts of the world. The multiregional hypothesis, on the other hand, proposes that modern humans evolved, at least in part, from independent hominid populations.[13]
Geneticists Lynn Jorde and Henry Harpending of the University of Utah proposed that the variation in human DNA is minute compared to that of other species. They also propose that during the Late Pleistocene, the human population was reduced to a small number of breeding pairs — no more than 10,000 and possibly as few as 1,000 — resulting in a very small residual gene pool. Various reasons for this hypothetical bottleneck have been postulated, one of those is the Toba catastrophe theory.
Human evolution is characterized by a number of important morphological, developmental, physiological and behavioral changes which have taken place since the split between the last common ancestor of Homo and Pan. The primary change, both in terms of chronology and in terms of it being the trait that defines the human subtribe the Hominina, was the evolution of a bipedal locomotor adaptation from an arboreal or semi-arboreal locomotor adaptation, with all its attendant adaptations, such as a valgus knee, low intermembral index (long legs relative to the arms) and reduced upper body strength.[citation needed] Following this was the evolution of a larger brain cavity and brain itself, which is typically 1,400 cm³ in modern humans; over twice that of a chimpanzee or gorilla. Other significant morphological changes included: the evolution of a power and precision grip;[citation needed] a reduced masticatory system; a reduction of the canine tooth; and the descent of the larynx and hyoid bone, making speech possible. With respect to development, the pattern of human postnatal brain growth differs from that of other apes (heterochrony), allowing for an extended period of social learning and language acquisition in juvenile humans. Physical anthropologists argue that a reorganization of the structure of the brain is more important than cranial expansion itself. One important physiological change in humans was the evolution of hidden estrus or concealed ovulation in females, which may have coincided with the evolution of important behavioral changes, such as pair bonding. Another significant behavioral change includes the development of material culture, or the (over time) increasingly wide variety of human-made objects which are used to manipulate humans' physical and social environments. How all these changes are related and what their role is in the evolution of complex social organization and culture are matters of ongoing debate.[14][15]
The closest living relatives of Homo sapiens are two distinct species of the genus Pan: the Bonobo (Pan paniscus) and the Common Chimpanzee (Pan troglodytes). These species share the same common ancestor. The main difference between them is the social organization: matriarchal for the Bonobo and patriarchal for the Common Chimpanzee. Full genome sequencing resulted in the conclusion that "after 6.5 million years of separate evolution, the differences between bonobo/chimpanzee and human are just 10 times greater than those between two unrelated people and 10 times less than those between rats and mice". In fact, 95 per cent of the DNA sequence is identical between the two Pan species and human.[8][9][10][11] It has been estimated that the human lineage diverged from that of chimpanzees about five million years ago, and from gorillas about eight million years ago. However, a hominid skull discovered in Chad in 2001, classified as Sahelanthropus tchadensis, is approximately seven million years old, which may indicate an earlier divergence.[12]
There are two prominent scientific theories of the origins of contemporary humans. They concern the relationship between modern humans and other hominids. The single-origin, or "out-of-Africa", hypothesis proposes that modern humans evolved in Africa and later migrated outwards to replace hominids in other parts of the world. The multiregional hypothesis, on the other hand, proposes that modern humans evolved, at least in part, from independent hominid populations.[13]
Geneticists Lynn Jorde and Henry Harpending of the University of Utah proposed that the variation in human DNA is minute compared to that of other species. They also propose that during the Late Pleistocene, the human population was reduced to a small number of breeding pairs — no more than 10,000 and possibly as few as 1,000 — resulting in a very small residual gene pool. Various reasons for this hypothetical bottleneck have been postulated, one of those is the Toba catastrophe theory.
Human evolution is characterized by a number of important morphological, developmental, physiological and behavioral changes which have taken place since the split between the last common ancestor of Homo and Pan. The primary change, both in terms of chronology and in terms of it being the trait that defines the human subtribe the Hominina, was the evolution of a bipedal locomotor adaptation from an arboreal or semi-arboreal locomotor adaptation, with all its attendant adaptations, such as a valgus knee, low intermembral index (long legs relative to the arms) and reduced upper body strength.[citation needed] Following this was the evolution of a larger brain cavity and brain itself, which is typically 1,400 cm³ in modern humans; over twice that of a chimpanzee or gorilla. Other significant morphological changes included: the evolution of a power and precision grip;[citation needed] a reduced masticatory system; a reduction of the canine tooth; and the descent of the larynx and hyoid bone, making speech possible. With respect to development, the pattern of human postnatal brain growth differs from that of other apes (heterochrony), allowing for an extended period of social learning and language acquisition in juvenile humans. Physical anthropologists argue that a reorganization of the structure of the brain is more important than cranial expansion itself. One important physiological change in humans was the evolution of hidden estrus or concealed ovulation in females, which may have coincided with the evolution of important behavioral changes, such as pair bonding. Another significant behavioral change includes the development of material culture, or the (over time) increasingly wide variety of human-made objects which are used to manipulate humans' physical and social environments. How all these changes are related and what their role is in the evolution of complex social organization and culture are matters of ongoing debate.[14][15]